The trip is not over, more to come, so stay tuned!
Go read previous installments of this series: Part 1, Part 2, and Part 3.
Also, go see Nick's here, here, here and here.
In tropical regions, a lot of times, good outcrops are along riverbanks. Puerto Rico is no exception and that is why the Rio G locality is so good, the exposure is kept “fresh” because of the nearly constant river erosion. Of course the drawback is that fossils are also lost if nobody visits these type of localities at least once a month or after big rainstorms (at least that’s what I used to do).
The picture above shows one such example. I spotted this bone fragment on the wall, very close to water level (a little more than half a meter). And, as you can see in the inset, the surface facing away from the rock looks freshly broken. I did collect the fossil, and unsuccessfully looked for additional fragments nearby.
Above is the picture of the fossil, in dorsal view (anterior towards the top), and the interpretative drawing. As it turns out the fossil was part of a sirenian skull. What was left of the fossil, is the anterior part of the frontals (Fr) and the nasals (N), the supraorbital processes of the frontals are missing. The convex frontal roof and shallow nasal incisure (the concave area between the frontals) are some characters that identify this fossil as belonging to a halitheriine dugongid. In fact, it is very similar to the same part of a much more complete skull that I collected from that locality several years ago (see below).
The figure above shows the more complete skull. A and B show a close-up dorsal view (anterior towards the top) of the area that was preserved in the fragmentary fossil. C is a dorsal view of the skull (anterior to the right) with the outline of the enlarged area in A. As you can see the frontal and its relationship with the nasals look much the same as the fragmentary fossil. It also displays a shallow nasal incisure at the posterior end of the mesorostral fossa (MRF) and convex frontal roof. In the more complete skull the premaxilla (Pmx) partially cover the nasals and the supraorbital processes (SOP) are preserved. (In A and C it is missing the left nasal process of the premaxilla which is loose and needs to be re-attached, but I was able to put it in the drawing).
I’m pretty certain that the fragmentary fossil belongs to the same species as the more complete specimen; it was a pity that part of it was lost previous to it being found. If you’re well acquainted with extinct sirenians, you can probably guess what genus this skull belongs to. So, go ahead and make a guess!
One of these localities (see picture below), I have visited at least since 2000, and up until very recently, we thought that the only formations present there were the Early Oligocene San Sebastián Formation and the overlying Lares Limestone. Now, thanks to new information regarding the stratigraphy of the Tertiary limestones of the north coast of Puerto Rico (Ortega Ariza, 2009), we know that in this locality, overlying the Lares Ls, there are also units of the Montebello Limestone. The age of the Lares Limestone and Montebello Limestone were designated as Late Oligocene – lower Early Miocene and upper Early Miocene, respectively (Seiglie & Moussa, 1984). New data, using strontium isotopes obtained from tubes of the pelecypod Kuphus incrassatus, seems to indicate, instead, that both formations span the Late Oligocene (Johnson et al., 2006; Ramírez et al., 2006; Ortega Ariza, 2009). If this is correct (more samples need to be run, hint, hint!!) I will like this outcrop even more (sorry, can't hide my love for the Oligocene)!!
Here's the one of my favorite outcrops, where the Lares and Montebello limestones are exposed. The arrow points to a sirenian fossil that is yet to be collected.
Of course, what I’ve been mostly searching in these localities are sirenian remains, but like I mentioned on that previous post, other vertebrates have also been collected. Interestingly, the best sirenian remains have been collected from the upper Lares Limestone, with a total (so far) of two skulls, and a set of nine articulated vertebrae (see picture below). There are more fossils but those will be collected in due time. As for the sirenian skulls, well, they are an important part of my thesis work and I will discuss them at some point in the future.
Some articulated sirenian vertebrae, these have already been collected. This is an earlier picture, there were three more vertebrae behind the one labeled Ca1, the ones anterior to L3 were collected earlier.
References
Johnson, C. C., W. R. Ramírez, L. R. Mark, S. Y. Hernandez, E. A. Barrow, M. Hegewald & J. Velez. 2006. Oligocene reef deposits linked to OPD site 999 with strontium isotope stratigraphy. Geological Society of America Abstracts with Program 38:557.
Ramírez, W. R., C. C. Johnson, M. Martínez, M. C. Torres & V. Ortiz. 2006. Strontium isotope stratigraphy from Kuphus incrassatus, Cenozoic limestones, Puerto Rico. Geological Society of America Abstracts with Program 38:90.
Seiglie, G. A. & M. T. Moussa. 1984. Late Oligocene-Pliocene trangressive-regressive cycles of sedimentation in northwestern Puerto Rico. American Association of Petroleum Geologist Memoir 36:89-95.